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Why We Love to Get Lost in Games: The Enduring Appeal of Metroidvanias
(www.nytimes.com)
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I count Dark Souls as a Metroidvania in my head, honestly.
But I think the actual defining feature is unlocking new abilities to reach new locations.
DS1 has you unlocking new areas that are interconnected with ones previously explored, but you don't really unlock new moves to get to a new place, it usually just happens after you beat a boss or buy an item.